Publications

156.    Huisman, J. 2017. Pedigree reconstruction from SNP data: parentage assignment, sibship clustering and beyond. Molecular Ecology Resources [tba] Click here

155.     Siepielski, A. M., M. B. Morrissey, M. Buoro, S. M. Carlson, C. M. Caruso, S. M. Clegg, T. Coulson, J. DiBattista, K. M. Gotanda, C. D. Francis, J. Hereford, J. G. Kingsolver, K. E. Augustine, L. E. B. Kruuk, R. A. Martin, B. C. Sheldon, N. Sletvold, E. I. Svensson, M. J. Wade, and A. D. C. MacColl. 2017. Precipitation drives global variation in natural selection. Science 355: 959-962. Click here

154.    Logan, C.J., L.E.B. Kruuk, R. Stanley, A.M. Thompson, and T.H. Clutton-Brock. 2016. Endocranial volume is heritable and is associated with longevity and fitness in a wild mammal. Royal Society Open Science 3: 160622. Click here

153.    Froy, H., C. A. Walling, J. M. Pemberton, T. H. Clutton-Brock, and L. E. B. Kruuk. 2016. Relative costs of offspring sex and offspring survival in a polygynous mammal. Biology Letters 12: 20160417. Click here

152.     Thackeray, S. J., Henrys, P. A., Hemming, D., Bell, J. R., Botham, M. S., Burthe, S., Helaouet, P., Johns, D. G., Jones, I. D., Leech, D. I., Mackay, E. B., Massimino, D., Atkinson, S., Bacon, P. J., Brereton, T. M., Carvalho, L., Clutton-Brock, T. H., Duck, C., Edwards, M., Elliott, J. M., Hall, S. J. G., Harrington, R., Pearce-Higgins, J W., Høye, T. T., Kruuk, L. E. B., Pemberton, J. M., Sparks, T. H., Thompson, P. M., White, I., Winfield, I. J. & Wanless, S. (2016) Phenological sensitivity to climate across taxa and trophic levels. Nature 535: 241-245. Click here

151.     Huisman, J., L. E. B. Kruuk, P. A. Ellis, T. Clutton-Brock, and J. M. Pemberton. 2016. Inbreeding depression across the lifespan in a wild mammal population. Proceedings of the National Acadamy of Sciences 113: 3585–3590. Click here 

150.     Moore, E. K., A. J. Britton, G. Iason, J. M. Pemberton and R. J. Pakeman. 2015. Landscape-scale vegetation patterns influence small-scale grazing impacts. Biological Conservation 192: 218-225. Click here

149.      Pavitt, A. T.,  C. A. Walling, E. Möstl, J. M. Pemberton, and L. E. Kruuk. 2015. Cortisol but not testosterone is repeatable and varies with reproductive effort in wild red deer stags. General and comparative endocrinology 222: 62-68. Click here

148.     Pavitt, A. T.,  C. A. Walling, J. M. Pemberton, and L. E. Kruuk. 2015. Heritability and cross-sex genetic correlations of early-life circulating testosterone levels in a wild mammal. Biology Letters 10: 20140685. Click here

147.     Lemaître, J.-F.,  J.-M. Gaillard, J. M. Pemberton, T. H. Clutton-Brock, and D. H. Nussey. 2014. Early life expenditure in sexual competition is associated with increased reproductive senescence in male red deer. Proceedings of the Royal Society B-Biological Sciences 281: 20140792. Click here

146.      Pérez-Barbería, F. J., E. I. Duff, M. J. Brewer, and F. E. Guinness. 2014. Evaluation of methods to age Scottish red deer: the balance between accuracy and practicality. Journal of Zoology 294: 180-189. Click here

145.      Walling, C. A., M. B. Morrissey, K. Foerster, T. H. Clutton-Brock, J. M. Pemberton, and L. E. B. Kruuk. 2014. A Multivariate Analysis of Genetic Constraints to Life History Evolution in a Wild Population of Red Deer. Genetics 198: 1735-1749. Click here

144.      Stopher, K. V., A. I. Bento, T. H. Clutton-Brock, J. M. Pemberton, and L. E. B. Kruuk. 2014. Multiple pathways mediate the effects of climate change on maternal reproductive traits in a red deer population. Ecology 95: 3124-3138. Click here

143.       Pavitt, A. T., C. A. Walling, A. S. McNeilly, J. M. Pemberton, and L. E. B. Kruuk. 2014. Variation in early-life testosterone within a wild population of red deer. Functional Ecology 28:1224-1234. Click here

142.       Andres, D., T. H. Clutton-Brock, L. E. B. Kruuk, J. M. Pemberton, K. V. Stopher and K. E. Ruckstuhl. 2013. Sex differences in the consequences of maternal loss in a long-lived mammal, the red deer (Cervus elaphus). Behavioral Ecology and Sociobiology 67:1249-1258. Click here

141.       Stopher, K. V., D. H. Nussey, T. H. Clutton‐Brock, F. Guinness, A. Morris and J. M. Pemberton. 2012. Re‐mating across years and intralineage polygyny are associated with greater than expected levels of inbreeding in wild red deer. Journal of evolutionary biology 25:2457-2469. Click here

140.       Morrissey, M. B., C. A. Walling, A. J. Wilson, J. M. Pemberton, T. H. Clutton-Brock, and L. E. Kruuk. 2012. Genetic analysis of life-history constraint and evolution in a wild ungulate population. The American Naturalist 179:E97-E114. Click here

139.       Stopher, K. V., C. A. Walling, A. Morris, F. E. Guinness, T. H. Clutton-Brock, J. M. Pemberton, and D. H. Nussey. 2012. Shared spatial effects on quantitative genetic parameters: accounting for spatial autocorrelation and home range overlap reduces estimates of heritability in wild red deer. Evolution 66:2411–2426. Click here

138.       Wilson, A. J., M. B. Morrissey, M. J. Adams, C. A. Walling, F. E. Guinness, J. M. Pemberton, T. H. Clutton-Brock, and L. E. B. Kruuk. 2011. Indirect genetics effects and evolutionary constraint: an analysis of social dominance in red deer, Cervus elaphus. Journal Of Evolutionary Biology 24:772-783.Click here

137.       Walling, C., D. Nussey, A. Morris, T. Clutton-Brock, L. Kruuk, and J. Pemberton. 2011. Inbreeding depression in red deer calves. BMC Evolutionary Biology 11:318.Click here

136.       Stopher, K., D. Nussey, T. Clutton-Brock, F. Guinness, A. Morris, and J. Pemberton. 2011. The red deer rut revisited: female excursions but no evidence females move to mate with preferred males. Behavioral Ecology 22:808-818.Click here

135.       Stevens, R. E., M. Balasse, and T. C. O'Connell. 2011. Intra-tooth oxygen isotope variation in a known population of red deer: Implications for past climate and seasonality reconstructions. Palaeogeography Palaeoclimatology Palaeoecology 301:64-74.Click here

134.       Moyes, K., D. H. Nussey, M. N. Clements, F. E. Guinness, A. Morris, S. Morris, J. M. Pemberton, L. E. B. Kruuk, and T. H. Clutton-Brock. 2011. Advancing breeding phenology in response to environmental change in a wild red deer population. Global Change Biology 17:2455-2469.Click here

133.        Moyes, K., B. Morgan, A. Morris, S. Morris, T. Clutton-Brock, and T. Coulson. 2011. Individual differences in reproductive costs examined using multi-state methods. Journal Of Animal Ecology 80:456-465. Click here

132.       Clements, M. N., T. H. Clutton-Brock, F. E. Guinness, J. M. Pemberton, and L. E. B. Kruuk. 2011. Variances and covariances of phenological traits in a wild mammal population. Evolution 65:788-801 Click here

131.       Clements, M. N., T. H. Clutton-Brock, S. D. Albon, J. M. Pemberton, and L. E. B. Kruuk. 2011. Gestation length variation in a wild ungulate. Functional Ecology 25:691-703. Click here

130.    Walling, C. A., J. M. Pemberton, J. D. Hadfield, and L. E. B. Kruuk. 2010. Comparing parentage inference software: reanalysis of a red deer pedigree. Molecular Ecology 19:1914-1928. Click here

129.    Clements, M. N., T. H. Clutton-Brock, S. D. Albon, J. M. Pemberton, and L. E. B. Kruuk. 2010. Getting the timing right: antler growth phenology and sexual selection in a wild red deer population. Oecologia 164:357-368. Click here

128.    Sisson, S. A. and Y. Fan. 2009. Towards automating model selection for a mark-recapture-recovery analysis. Journal of the Royal Statistical Society Series C-Applied Statistics 58:247-266.Click here

127.    Nussey, D. H., L. E. B. Kruuk, A. Morris, M. N. Clements, J. M. Pemberton, and T. H. Clutton-Brock. 2009. Inter- and Intrasexual Variation in Aging Patterns across Reproductive Traits in a Wild Red Deer Population. American Naturalist 174:342-357.Click here

126.    Moyes, K., B. J. T. Morgan, A. Morris, S. J. Morris, T. H. Clutton-Brock, and T. Coulson. 2009. Exploring individual quality in a wild population of red deer. Journal Of Animal Ecology 78:406-413.Click here

125.    Stopher, K. V., T. H. Clutton-Brock, J. M. Pemberton, and T. Coulson. 2008. Individual differences, density dependence and offspring birth traits in a population of red deer. Proceedings Of The Royal Society Of London Series B-Biological Sciences 275:2137–2145.Click here

124.    Nussey, D. H., A. J. Wilson, A. Morris, J. Pemberton, T. Clutton-Brock, and L. E. B. Kruuk. 2008. Testing for genetic trade-offs between early- and late-life reproduction in a wild red deer population. Proceedings of the Royal Society B-Biological Sciences 275:745-750.Click here

123.    Nussey, D. H., T. Coulson, M. Festa-Bianchet, and J. M. Gaillard. 2008. Measuring senescence in wild animal populations: towards a longitudinal approach. Functional Ecology 22:393-406. Click here

122.    McLoughlin, P. D., T. Coulson, and T. Clutton-Brock. 2008. Cross-generational effects of habitat and density on life history in red deer. Ecology 89:3317-3326.Click here

121.    Jones, O. R., J. M. Gaillard, S. Tuljapurkar, J. S. Alho, K. B. Armitage, P. H. Becker, P. Bize, J. Brommer, A. Charmantier, M. Charpentier, T. Clutton-Brock, F. S. Dobson, M. Festa-Bianchet, L. Gustafsson, H. Jensen, C. G. Jones, B. G. Lillandt, R. McCleery, J. Merila, P. Neuhaus, M. A. C. Nicoll, K. Norris, M. K. Oli, J. Pemberton, H. Pietiainen, T. H. Ringsby, A. Roulin, B. E. Saether, J. M. Setchell, B. C. Sheldon, P. M. Thompson, H. Weimerskirch, E. J. Wickings, and T. Coulson. 2008. Senescence rates are determined by ranking on the fast-slow life-history continuum. Ecology Letters 11:664-673.Click here

120.    Hadfield, J. D. 2008. Estimating evolutionary parameters when viability selection is operating. Proceedings Of The Royal Society Of London Series B-Biological Sciences 275:723-734.Click here

119.    Fortin, D., D. W. Morris, and P. D. McLoughlin. 2008. Habitat selection and the evolution of specialists in heterogeneous environments. Israel Journal of Ecology & Evolution 54:311-328.Click here

118.    Coulson, T. and S. Tuljapurkar. 2008. The dynamics of a quantitative trait in an age-structured population living in a variable environment. American Naturalist 172:E599-E612.Click here

117.    Charlton, B. D., K. McComb, and D. Reby. 2008. Free-ranging red deer hinds show greater attentiveness to roars with formant frequencies typical of young males. Ethology 114:1023-1031.Click here

116.    Wilson, A. J., D. H. Nussey, J. M. Pemberton, J. G. Pilkington, A. Morris, F. Pelletier, T. H. Clutton-Brock, and L. E. B. Kruuk. 2007. Evidence for a genetic basis of aging in two wild vertebrate populations. Current Biology 17:2136-2142.Click here

115.    Slate, J. and J. M. Pemberton. 2007. Admixture and patterns of linkage disequilibrium in a free-living vertebrate population. Journal Of Evolutionary Biology 20:1415-1427.Click here

114.    Sims, M., D. A. Elston, A. Larkham, D. H. Nussey, and S. D. Albon. 2007. Identifying when weather influences life-history traits of grazing herbivores. Journal Of Animal Ecology 76:761-770.Click here

113.    Nussey, D. H., B. Metherell, K. Moyes, A. Donald, F. E. Guinness, and T. H. Clutton-Brock. 2007. The relationship between tooth wear, habitat quality and late-life reproduction in a wild red deer population. Journal Of Animal Ecology 76:402-412.Click here

112.    Nussey, D. H., L. E. B. Kruuk, A. Morris, and T. H. Clutton-Brock. 2007. Environmental conditions in early life influence ageing rates in a wild population of red deer. Current Biology 17:R1000-R1001.Click here

111.    Nilsen, E. B., E. J. Milner-Gulland, L. Schofield, A. Mysterud, N. C. Stenseth, and T. Coulson. 2007. Wolf reintroduction to Scotland: public attitudes and consequences for red deer management. Proceedings of the Royal Society B-Biological Sciences 274:995-1002.Click here

110.    Kruuk, L. E. B. and J. D. Hadfield. 2007. How to separate genetic and environmental causes of similarity between relatives. Journal Of Evolutionary Biology 20:1890-1903.Click here

109.    Kalinowski, S. T., M. L. Taper, and T. C. Marshall. 2007. Revising how the computer program CERVUS accommodates genotyping error increases success in paternity assignment. Molecular Ecology 16:1099-1106.Click here

108.    Hone, J. and T. H. Clutton-Brock. 2007. Climate, food, density and wildlife population growth rate. Journal Of Animal Ecology 76:361-367.Click here

107.    Foerster, K., T. Coulson, B. C. Sheldon, J. M. Pemberton, T. H. Clutton-Brock, and L. E. B. Kruuk. 2007. Sexually antagonistic genetic variation for fitness in red deer. Nature 447:1107-1109.Click here

106.    Stevens, R. E., A. M. Lister, and R. E. M. Hedges. 2006. Predicting diet, trophic level and palaeoecology from bone stable isotope analysis: a comparative study of five red deer populations. Oecologia 149:12-21.Click here

105.    Nussey, D. H., J. Pemberton, A. Donald, and L. E. B. Kruuk. 2006. Genetic consequences of human management in an introduced island population of red deer (Cervus elaphus). Heredity 97:56-65.Click here

104.    Nussey, D. H., L. E. B. Kruuk, A. Donald, M. Fowlie, and T. H. Clutton-Brock. 2006. The rate of senescence in maternal performance increases with early-life fecundity in red deer. Ecology Letters 9:1342-1350.Click here

103.    Moyes, K., T. Coulson, B. J. T. Morgan, A. Donald, S. J. Morris, and T. H. Clutton-Brock. 2006. Cumulative reproduction and survival costs in female red deer. Oikos 115:241-252.Click here

102.    McLoughlin, P. D., M. S. Boyce, T. Coulson, and T. Clutton-Brock. 2006. Lifetime reproductive success and density-dependent, multi-variable resource selection. Proceedings of the Royal Society B-Biological Sciences 273:1449-1454.Click here

101.    Lande, R., S. Engen, B. E. Saether, and T. Coulson. 2006. Estimating density dependence from time series of population age structure. American Naturalist 168:76-87.Click here

100.    Csillery, K., T. Johnson, D. Beraldi, T. Clutton-Brock, D. Coltman, B. Hansson, G. Spong, and J. M. Pemberton. 2006. Performance of marker-based relatedness estimators in natural populations of outbred vertebrates. Genetics 173:2091-2101.Click here

99.    Reby, D., K. McComb, B. Cargnelutti, C. Darwin, W. T. Fitch, and T. Clutton-Brock. 2005. Red deer stags use formants as assessment cues during intrasexual agonistic interactions. Proceedings of the Royal Society B-Biological Sciences 272:941-947.Click here

98.    Nussey, D. H., D. W. Coltman, T. Coulson, L. E. B. Kruuk, A. Donald, S. J. Morris, T. H. Clutton-Brocks, and J. Pemberton. 2005. Rapidly declining fine-scale spatial genetic structure in female red deer. Molecular Ecology 14:3395-3405.Click here

97.    Nussey, D. H., T. H. Clutton-Brock, D. A. Elston, S. D. Albon, and L. E. B. Kruuk. 2005. Phenotypic plasticity in a maternal trait in red deer. Journal Of Animal Ecology 74:387-396.Click here

96.    Nussey, D. H., T. H. Clutton-Brock, S. D. Albon, J. Pemberton, and L. E. B. Kruuk. 2005. Constraints on plastic responses to climate variation in red deer. Biology Letters 1:457-460.Click here

95.    Coulson, T., J. M. Gaillard, and M. Festa-Bianchet. 2005. Decomposing the variation in population growth into contributions from multiple demographic rates. Journal Of Animal Ecology 74:789-801.Click here

94.    Milner-Gulland, E. J., T. Coulson, and T. H. Clutton-Brock. 2004. Sex differences and data quality as determinants of income from hunting red deer Cervus elaphus. Wildlife Biology 10:187-201.

93.    Coulson, T., F. Guinness, J. Pemberton, and T. Clutton-Brock. 2004. The demographic consequences of releasing a population of red deer from culling. Ecology 85:411-422.Click here

92.    Catchpole, E. A., Y. Fan, B. J. T. Morgan, T. H. Clutton-Brock, and T. Coulson. 2004. Sexual dimorphism, survival and dispersal in red deer. Journal of Agricultural Biological and Environmental Statistics 9:1-26.Click here

91.    Reby, D. and K. McComb. 2003. Anatomical constraints generate honesty: acoustic cues to age and weight in the roars of red deer stags. Animal Behaviour 65:519-530.Click here

90.    Kruuk, L. E. B., J. Slate, J. M. Pemberton, and T. H. Clutton-Brock. 2003. Fluctuating asymmetry in a secondary sexual trait: no associations with individual fitness, environmental stress or inbreeding, and no heritability. Journal Of Evolutionary Biology 16:101-113.Click here

89.    Coulson, T., L. E. B. Kruuk, G. Tavecchia, J. M. Pemberton, and T. H. Clutton-Brock. 2003. Estimating selection on neonatal traits in red deer using elasticity path analysis. Evolution 57:2879-2892.Click here

88.    Virtanen, R., G. R. Edwards, and M. J. Crawley. 2002. Red deer management and vegetation on the Isle of Rum. Journal Of Applied Ecology 39:572-583.Click here

87.    Slate, J., P. M. Visscher, S. MacGregor, D. Stevens, M. L. Tate, and J. M. Pemberton. 2002. A genome scan for quantitative trait loci in a wild population of red deer (Cervus elaphus). Genetics 162:1863-1873.Click here

86.    Slate, J. and J. M. Pemberton. 2002. Comparing molecular measures for detecting inbreeding depression. Journal Of Evolutionary Biology 15:20-31.Click here

85.    Marshall, T. C., D. W. Coltman, J. M. Pemberton, J. Slate, J. A. Spalton, F. E. Guinness, J. A. Smith, J. G. Pilkington, and T. H. Clutton-Brock. 2002. Estimating the prevalence of inbreeding from incomplete pedigrees. Proceedings Of The Royal Society Of London Series B-Biological Sciences 269:1533-1539.Click here

84.    Kruuk, L. E. B., J. Slate, J. M. Pemberton, S. Brotherstone, F. Guinness, and T. Clutton-Brock. 2002. Antler size in red deer: Heritability and selection but no evolution. Evolution 56:1683-1695.Click here

83.    Clutton-Brock, T. H., T. N. Coulson, E. J. Milner-Gulland, D. Thomson, and H. M. Armstrong. 2002. Sex differences in emigration and mortality affect optimal management of deer populations. Nature 415:633-637.Click here

82.    Clutton-Brock, T. H. and T. Coulson. 2002. Comparative ungulate dynamics: the devil is in the detail. Philosophical Transactions Of The Royal Society Of London Series B-Biological Sciences 357:1285-1298.Click here

81.    Schmidt, K. T., A. Stien, S. D. Albon, and F. E. Guinness. 2001. Antler length of yearling red deer is determined by population density, weather and early life-history. Oecologia 127:191-197.Click here

80.    Conradt, L., I. J. Gordon, T. H. Clutton-Brock, D. Thomson, and F. E. Guinness. 2001. Could the indirect competition hypothesis explain inter-sexual site segregation in red deer (Cervus elaphus L.)? Journal Of Zoology 254:185-193.Click here

79.    Slate, J., T. Marshall, and J. Pemberton. 2000. A retrospective assessment of the accuracy of the paternity inference program CERVUS. Molecular Ecology 9:801-808.Click here

78.    Slate, J., L. E. B. Kruuk, T. C. Marshall, J. M. Pemberton, and T. H. Clutton-Brock. 2000. Inbreeding depression influences lifetime breeding success in a wild population of red deer (Cervus elaphus). Proceedings Of The Royal Society Of London Series B-Biological Sciences 267:1657-1662.Click here

77.    Milner-Gulland, E. J., T. N. Coulson, and T. H. Clutton-Brock. 2000. On harvesting a structured ungulate population. Oikos 88:592-602.Click here

76.    Kruuk, L. E. B., T. H. Clutton-Brock, J. Slate, J. M. Pemberton, S. Brotherstone, and F. E. Guinness. 2000. Heritability of fitness in a wild mammal population. Proceedings Of The National Academy Of Sciences Of The United States Of America 97:698-703.Click here

75.    Coulson, T., E. J. Milner-Gulland, and T. Clutton-Brock. 2000. The relative roles of density and climatic variation on population dynamics and fecundity rates in three contrasting ungulate species. Proceedings Of The Royal Society Of London Series B-Biological Sciences 267:1771-1779.Click here

74.    Conradt, L., T. H. Clutton-Brock, and F. E. Guinness. 2000. Sex differences in weather sensitivity can cause habitat segregation: red deer as an example. Animal Behaviour 59:1049-1060.Click here

73.    Conradt, L. 2000. Use of a seaweed habitat by red deer (Cervus elaphus L.). Journal Of Zoology 250:541-549.Click here

72.    Albon, S. D., T. N. Coulson, D. Brown, F. E. Guinness, J. M. Pemberton, and T. H. Clutton-Brock. 2000. Temporal changes in key factors and key age groups influencing the population dynamics of female red deer. Journal Of Animal Ecology 69:1099-1110.Click here

71.    Slate, J., J. M. Pemberton, and P. M. Visscher. 1999. Power to detect QTL in a free-living polygynous ruminant population. Heredity 83:327-336.Click here

70.    Pemberton, J. M., D. W. Coltman, T. N. Coulson, and J. Slate. 1999. Using microsatellites to measure the fitness consequences of inbreeding and outbreeding.in D. Goldstein and C. Schlotterer, editors. Microsatellites: Evolution and Applications. Oxford University Press, Oxford.

69.    Kruuk, L. E. B., T. H. Clutton-Brock, K. E. Rose, and F. E. Guinness. 1999. Early determinants of lifetime reproductive success differ between the sexes in red deer. Proceedings Of The Royal Society Of London Series B-Biological Sciences 266:1655-1661.Click here

68.    Kruuk, L. E. B., T. H. Clutton-Brock, S. D. Albon, J. M. Pemberton, and F. E. Guinness. 1999. Population density affects sex ratio variation in red deer. Nature 399:459-461.Click here

67.    Coulson, T., S. Albon, J. Slate, and J. Pemberton. 1999. Microsatellite loci reveal sex-dependent responses to inbreeding and outbreeding in red deer calves. Evolution 53:1951-1960.Click here

66.    Conradt, L., T. H. Clutton-Brock, and D. Thomson. 1999. Habitat segregation in ungulates: are males forced into suboptimal foraging habitats through indirect competition by females? Oecologia 119:367-377.Click here

65.    Conradt, L., T. H. Clutton-Brock, and F. E. Guinness. 1999. The relationship between habitat choice and lifetime reproductive success in female red deer. Oecologia 120:218-224.Click here

64.    Conradt, L. 1999. Social segregation is not a consequence of habitat segregation in red deer and feral soay sheep. Animal Behaviour 57:1151-1157.Click here

63.    Slate, J., D. W. Coltman, S. J. Goodman, I. MacLean, J. M. Pemberton, and J. L. Williams. 1998. Bovine microsatellite loci are highly conserved in red deer (Cervus elaphus), sika deer (Cervus nippon) and Soay sheep (Ovis aries). Animal Genetics 29:307-315.Click here

62.    Rose, K. E., T. H. Clutton-Brock, and F. E. Guinness. 1998. Cohort variation in male survival and lifetime breeding success in red deer. Journal Of Animal Ecology 67:979-986.Click here

61.    Pemberton, J. M., J. A. Smith, T. N. Coulson, T. C. Marshall, J. Slate, S. Paterson, S. D. Albon, and T. H. Clutton-Brock. 1998. The maintenance of genetic polymorphism in small island populations: large mammals in the Hebrides.in P. R. Grant, editor. Evolution and Islands. Oxford University Press, Oxford.

60.    Marshall, T. C., J. Slate, L. E. B. Kruuk, and J. M. Pemberton. 1998. Statistical confidence for likelihood-based paternity inference in natural populations. Molecular Ecology 7:639-655.Click here

59.    Coulson, T. N., J. M. Pemberton, S. D. Albon, M. Beaumont, T. C. Marshall, J. Slate, F. E. Guinness, and T. H. Clutton-Brock. 1998. Microsatellites reveal heterosis in red deer. Proceedings Of The Royal Society Of London Series B-Biological Sciences 265:489-495.Click here

58.    Coulson, T. N., S. D. Albon, J. M. Pemberton, J. Slate, F. E. Guinness, and T. H. Clutton-Brock. 1998. Genotype by environment interactions in winter survival in red deer. Journal Of Animal Ecology 67:434-445.Click here

57.    Conradt, L. 1998. Measuring the degree of sexual segregation in group-living animals. Journal Of Animal Ecology 67:217-226.Click here

56.    Conradt, L. 1998. Could asynchrony in activity between the sexes cause intersexual social segregation in ruminants? Proceedings Of The Royal Society Of London Series B-Biological Sciences 265:1359-1363.Click here

55.    Flint, A. P. F., S. D. Albon, A. S. I. Loudon, and J. H.N. 1997. Behavioral dominance and corpus luteum function in red deer Cervus elaphus. Hormones and Behaviour 31:296-304.Click here

54.    Flint, A. P. F., S. D. Albon, and S. I. Jafar. 1997. Blastocyst development and conceptus sex selection in red deer Cervus elaphus: Studies of a free-living population on the Isle of Rum. General and Comparative Endocrinology 106:374-383.Click here

53.    Coulson, T., S. Albon, F. Guinness, J. Pemberton, and T. CluttonBrock. 1997. Population substructure, local density, and calf winter survival in red deer (Cervus elaphus). Ecology 78:852-863.Click here

52.    Clutton-Brock, T. H., K. E. Rose, and F. E. Guinness. 1997. Density-related changes in sexual selection in red deer. Proceedings Of The Royal Society Of London Series B-Biological Sciences 264:1509-1516.Click here

51.    Clutton-Brock, T. H., A. W. Illius, K. Wilson, B. T. Grenfell, A. D. C. MacColl, and S. D. Albon. 1997. Stability and instability in ungulate populations: An empirical analysis. American Naturalist 149:195-219.Click here

50.  Pemberton, J. M., J. A. Smith, T. N. Coulson, T. C. Marshall, J. Slate, S. Paterson, S. D. Albon, and T. H. Clutton-Brock. 1996. The maintenance of genetic polymorphism in small island populations: Large mammals in the Hebrides. Philosophical Transactions Of The Royal Society Of London Series B-Biological Sciences 351:745-752.Click here

49.  Langvatn, R., S. D. Albon, T. Burkey, and T. H. Clutton-Brock. 1996. Climate, plant phenology and variation in age of first reproduction in a temperate herbivore. Journal Of Animal Ecology 65:653-670.Click here

48.  Pemberton, J. M., J. Slate, D. R. Bancroft, and J. A. Barrett. 1995. Nonamplifying Alleles At Microsatellite Loci - A Caution For Parentage And Population Studies. Molecular Ecology 4:249-252.Click here

47.  Clutton-Brock, T. H. and G. A. Parker. 1995. Sexual Coercion in Animal Societies. Animal Behaviour 49:1345-1365.Click here

46.  Benton, T. G., A. Grant, and T. H. Cluttonbrock. 1995. Does Environmental Stochasticity Matter - Analysis of Red Deer Life-Histories on Rum. Evolutionary Ecology 9:559-574.Click here

45.  Clutton-Brock, T. H. and M. E. Lonergan. 1994. Culling Regimes And Sex Ratio Biases In Highland Red Deer. Journal Of Applied Ecology 31:521-527.Click here

44.  Pemberton, J. M., S. D. Albon, F. E. Guinness, T. H. Clutton-Brock, and G. A. Dover. 1992. Behavioral Estimates Of Male Mating Success Tested By DNA Fingerprinting In A Polygynous Mammal. Behavioral Ecology 3:66-75.Click here

43.  Albon, S. D., H. J. Staines, F. E. Guinness, and T. H. Clutton-Brock. 1992. Density-Dependent Changes In The Spacing Behavior Of Female Kin In Red Deer. Journal Of Animal Ecology 61:131-137.Click here

42.  Pemberton, J. M., S. D. Albon, F. E. Guinness, and T. H. Clutton-Brock. 1991. Countervailing Selection In Different Fitness Components In Female Red Deer. Evolution 45:93-103.Click here

41.  Gomendio, M., T. H. Clutton-Brock, S. D. Albon, F. E. Guinness, and M. J. Simpson. 1990. Mammalian Sex Ratios And Variation In Costs Of Rearing Sons And Daughters. Nature 343:261-263.Click here

40.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1989. Fitness Costs of Gestation and Lactation in Wild Mammals. Nature 337:260-262.Click here

39.  Pemberton, J. M., S. D. Albon, F. E. Guinness, T. H. Clutton-Brock, and R. J. Berry. 1988. Genetic Variation And Juvenile Survival In Red Deer. Evolution 42:921-934.Click here

38.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1988. Reproductive success in male and female red deer. Pages 325-343 in T. H. Clutton-Brock, editor. Reproductive success. University of Chicago Press, Chicago.

37.  McComb, K. 1987. Roaring By Red Deer Stags Advances The Date Of Estrus In Hinds. Nature 330:648-649.Click here

36.  Clutton-Brock, T. H., M. Major, S. D. Albon, and F. E. Guinness. 1987. Early Development And Population Dynamics In Red Deer .1. Density-Dependent Effects On Juvenile Survival. Journal Of Animal Ecology 56:53-67.Click here

35.  Clutton-Brock, T. H., G. R. Iason, and F. E. Guinness. 1987. Sexual Segregation And Density-Related Changes In Habitat Use In Male And Female Red Deer (Cervus-Elaphus). Journal Of Zoology 211:275-289.Click here

34.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1987. Interactions Between Population-Density And Maternal Characteristics Affecting Fecundity And Juvenile Survival In Red Deer. Journal Of Animal Ecology 56:857-871.Click here

33.  Albon, S. D., T. H. Clutton-Brock, and F. E. Guinness. 1987. Early Development And Population Dynamics In Red Deer .2. Density-Independent Effects And Cohort Variation. Journal Of Animal Ecology 56:69-81.Click here

32.  Thouless, C. R. and F. E. Guinness. 1986. Conflict Between Red Deer Hinds - The Winner Always Wins. Animal Behaviour 34:1166-1171.Click here

31.  Iason, G. R., C. D. Duck, and T. H. Clutton-Brock. 1986. Grazing And Reproductive Success Of Red Deer - The Effect Of Local Enrichment By Gull Colonies. Journal Of Animal Ecology 55:507-515.Click here

30.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1986. Great Expectations - Dominance, Breeding Success And Offspring Sex Ratios In Red Deer. Animal Behaviour 34:460-471.Click here

29.  Albon, S. D., B. Mitchell, B. J. Huby, and D. Brown. 1986. Fertility In Female Red Deer (Cervus elaphus) - The Effects Of Body-Composition, Age And Reproductive Status. Journal Of Zoology 209:447-460.Click here

28.  Iason, G. R. and F. E. Guinness. 1985. Synchrony Of Estrus And Conception In Red Deer (Cervus elaphus L). Animal Behaviour 33:1169-1174.Click here

27.  Clutton-Brock, T. H., M. Major, and F. E. Guinness. 1985. Population Regulation In Male And Female Red Deer. Journal Of Animal Ecology 54:831-846.Click here

26.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1985. Parental Investment And Sex-Differences In Juvenile Mortality In Birds And Mammals. Nature 313:131-133.Click here

25.  Clutton-Brock, T. H. 1985. Reproductive Success In Red Deer. Scientific American 252:86.Click here

24.  Clutton-Brock, T. H., F. E. Guinness, and S. D. Albon. 1984. Individuals And Populations - The Effects Of Social-Behavior On Population-Dynamics In Deer. Proceedings Of The Royal Society Of Edinburgh Section B-Biological Sciences 82:275-290.Click here

23.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1984. Maternal Dominance, Breeding Success And Birth Sex Ratios In Red Deer. Nature 308:358-360.Click here

22.  Clutton-Brock, T. H. 1984. Reproductive Effort And Terminal Investment In Iteroparous Animals. American Naturalist 123:212-229.Click here

21.  Clutton-Brock, T. H., F. E. Guinness, and S. D. Albon. 1983. The Costs Of Reproduction To Red Deer Hinds. Journal Of Animal Ecology 52:367-383.Click here

20.  Clutton-Brock, T. H. and S. D. Albon. 1983. Climatic Variation And Body Weight Of Red Deer. Journal Of Wildlife Management 47:1197-1201.Click here

19.  Albon, S. D., B. Mitchell, and B. W. Staines. 1983. Fertility And Body Weight In Female Red Deer - A Density-Dependent Relationship. Journal Of Animal Ecology 52:969-980.Click here

18.  Albon, S. D., F. E. Guinness, and T. H. Clutton-Brock. 1983. The Influence Of Climatic Variation On The Birth Weights Of Red Deer (Cervus elaphus). Journal Of Zoology 200:295-298.Click here

17.  Clutton-Brock, T. H., G. R. Iason, S. D. Albon, and F. E. Guinness. 1982. Effects Of Lactation On Feeding Behavior And Habitat Use In Wild Red Deer Hinds. Journal Of Zoology 198:227-236.Click here

16.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1982. Competition between Female Relatives in a Matrilocal Mammal. Nature 300:178-180.Click here

15.  Clutton-Brock, T. H. and S. D. Albon. 1982. Winter Mortality In Red Deer (Cervus elaphus). Journal Of Zoology 198:515-519.

14.  Clutton-Brock, T. H. 1982. The Functions Of Antlers. Behaviour 79:108-125.Click here

13.  Clutton-Brock, T. 1982. The Red Deer Of Rhum. Natural History 91:42.

12.  Clutton-Brock, T. H., S. D. Albon, and F. E. Guinness. 1981. Parental Investment In Male And Female Offspring In Polygynous Mammals. Nature 289:487-489.Click here

11.  Gibson, R. M. and F. E. Guinness. 1980. Differential Reproduction Among Red Deer (Cervus elaphus) Stags On Rhum. Journal Of Animal Ecology 49:199-208.The Red Deer Of Rhum. Natural History 91:42.

10.  Gibson, R. M. and F. E. Guinness. 1980. Behavioral-Factors Affecting Male Reproductive Success in Red Deer (Cervus-Elaphus). Animal Behaviour 28:1163-1174.

9.  Guinness, F. E., M. J. Hall, and R. A. Cockerill. 1979. Mother-Offspring Association In Red Deer (Cervus elaphus L) On Rhum. Animal Behaviour 27:536-544.Click here

8.  Clutton-Brock, T. H., S. D. Albon, R. M. Gibson, and F. E. Guinness. 1979. Logical Stag - Adaptive Aspects Of Fighting In Red Deer (Cervus elaphus L). Animal Behaviour 27:211-225.Click here

7.  Clutton-Brock, T. H. and S. D. Albon. 1979. Roaring Of Red Deer And The Evolution Of Honest Advertisement. Behaviour 69:145-170.Click here

6.  Guinness, F. E., R. M. Gibson, and T. H. Clutton-Brock. 1978. Calving Times Of Red Deer (Cervus-Elaphus) On Rhum. Journal Of Zoology 185:105-114.Click here

5.  Guinness, F. E., T. H. Clutton-Brock, and S. D. Albon. 1978. Factors Affecting Calf Mortality In Red Deer (Cervus elaphus). Journal Of Animal Ecology 47:817-832.Click here

4.  Guinness, F. E., S. D. Albon, and T. H. Clutton-Brock. 1978. Factors Affecting Reproduction In Red Deer (Cervus-Elaphus) Hinds On Rhum. Journal Of Reproduction And Fertility 54:325-334.Click here

3.  Clutton-Brock, T. H. and F. E. Guinness. 1975. Behavior Of Red Deer (Cervus elaphus L) At Calving Time. Behaviour 55:287-300.Click here

2.  Lincoln, G. A. 1972. Role of Antlers in Behavior of Red Deer. Journal of Experimental Zoology 182:233-249.Click here

1.  Guinness, F., G. A. Lincoln, and R. V. Short. 1971. Reproductive Cycle of Female Red Deer, Cervus-Elaphus L. Journal Of Reproduction And Fertility 27:427-438.Click here